b. enoplocampa

Culture

Endangered Status

Endangered B1ab(iii),B2ab(iii).

Original Botanical Description or Link to

Perennial, monoecious, rhizomatous herb, up to ca. 40 cm tall. Stem rhizomatous, up to ca. 10 cm long, internodes strongly compressed, densely covered by branched, erect emergences to 1.9 cm long. Leaves alternate; stipules persistent, broadly ovate, asymmetric, 7–10 × 14–16 mm, margin entire, apex rounded, midrib abaxially prominent, forming a subapically emerging, branched bristle 5–15 mm long; petioles 22–37 cm long, terete, green with numerous small whitish dots or stripes, sparsely hairy with multicellular hairs to ca. 2 mm long and microscopic glandular hairs; lamina basifixed, 15–27 × 11–20 cm, broadly ovate, elliptic or suborbicular, asymmetric, margin dentate and denticulate between the larger teeth, often shallowly lobed (to ca. 30% of the lamina width), teeth bristlepointed, base cordate, lobes not overlapping or sometimes slightly overlapping, apex acute, adaxially green, sparsely bristly hairy on veins and lamina, abaxially pale green, sparsely hairy on veins only, venation palmate, primary veins 7–9, actinodromus, secondary veins craspedodromus. Inflorescences: bisexual, protandrous, axillary, dichasialmonochasial cymes, with 3–4 basal dichasial branchings and 1–3 distal monochasial branchings; peduncle 14–27.5 cm long, pale green, sparsely hairy; bracts anisophyllus, ovate to elliptic, 3–5 × 2–3 mm, minute in more distal inflorescence part, sparsely hairy, semi-persistent. Male flowers: pedicels 8–13 mm long, glabrescent, pale greenwhitish; tepals 4, unequal, outer tepals 2, much larger than inner, orbicular to broadly ovate, 8–15 × 10–16 mm, white, margin entire, apex rounded, glabrous, inner tepals 2, obovate to elliptic, 6–10 × 4–6 mm, margin entire, apex obtuse and shortly cuspidate, white; androecium yellow, symmetric, globose; stamens ca. 69–81, free filaments up to ca. 0.5 mm long, fused at the base to a short column, anthers up to ca. 0.5 mm long, dehiscing through laterally positioned slits > ½ as long as the anthers. Female flowers: pedicel 8 mm long, glabrous, whitish; tepals 3, unequal, 2 outer broadly ovate, 7 × 10 mm, margin entire, apex rounded, 1 inner tepal elliptic, 8 × 3 mm, apex acute; ovary (excluding the wings) 6 × 3 mm, ellipsoid, white, turning green at maturity (transition to fruiting), locules 3, placentation axile, placentae entire, wings 3, equal, base convex to subtruncate, apex rounded, cuneate, or subtruncate, widest point 4 mm (at the middle to basal part), style ca. 5 mm long, fused at base, 3–branched, each stylodium bifurcate in the stigmatic region, stigmatic surface a spirally twisted papillose band, yellow. Fruits: dry capsule; pedicels 9–20 mm long; seed-bearing part 7–10 × 5–7 mm (excluding the wings), ellipsoid, wing shape as for ovary, widest point up to 8 mm (middle to basally). Seeds barrel-shaped, ca. 0.3 mm long.