B. daunhitam

Culture

Endangered Status

Vulnerable

Original Botanical Description or Link to

Low herb, terrestrial, perennial. Stem erect to 20 cm high, 4–8 mm in diameter, old stem yellowish and slightly woody, young part red to brownish, fleshy and succulent, surface glabrous, internodes 1–3 cm long. Stipule red to brownish or purplish, ovate-triangular, 8– 10 mm long, 4–8 mm wide, keeled, margin entire, short cusp ca.1 mm long. Petiole terete, 5–20 mm long, 2–5 mm wide, brownish red, glabrous. Leaves up to 10 on branched stems, alternate, simple, lamina elliptic-ovate to oblong-ovate, 13–20 cm long (basal lobes included), 7–10 cm wide, broad side 3–4.5 cm wide, strongly asymmetric with a well-developed basal lobe on one side giving a cordate appearance, base unequal, basal lobes cordate, 2–3 cm long, overlapping when mature, margin denticulate, apex shortly acuminate; adaxially shiny, dark olive to solid black at maturity, immature stage dark bluish green, young leaves brown with a narrow red rim at margin, adaxial surface strongly bullate, giving a bubble-like appearance; abaxially light red to crimson, glabrescent; venation palmate-pinnate, midrib distinguishable, with ca. 3 major lateral veins on each side, other 4–5 primary veins branching dichotomously. Inflorescence bisexual, protogynous, terminal or axillary, erect, 10–25 cm long; basal branch with a pair of female flowers placed on short peduncle ca. 1 cm; upper distal branching dichasially cymose to 15 cm long, zig-zag, peduncle to 8 cm long, cymes branching to 4 orders, crimson to purplish or brownish, glabrous or with sparse white scales; bract at base of inflorescence narrowly lanceolate ca. 4 × 1 mm, red to brown, margin entire, apex cuspate; bracts at apex smaller than bract ca. 2 × 1 mm. Staminate flower: pedicel 5–12 mm long, glabrous; tepals 2, pink-red, elliptic, glabrous, 6–8 × 3–5 mm, margin entire; androecium symmetric, stamens 20–30, yellow; filaments 1–2 mm long, subequal, slightly fused at base; anthers obovate, 0.8–1 × 0.6–0.8 mm. Pistillate flower: pedicel 10–16 mm long, 1–2 mm in diameter, pink-red, glabrous; ovary 3-locular, 10–14 × 12–16 mm, light red to crimson, glabrous, placenta bilamellate; tepals 5, pink to light red, glabrous, base cuneate to rounded, apex acute to obtuse or slightly pointed, outer tepals 2, broadly elliptic, 8–10 × 4–8 mm, inner tepals 3, narrowly elliptic, 7–9 × 3–4.5 mm; styles 3, golden yellow, bifid, ca. 4 mm long, stigmas broadly Y-shaped, forming a spiral band, papillose. Capsule nodding, 12– 16 × 14–18 mm (wings included), pink to crimson, glabrous, wings 3, subequal, nearly parallel but rounded or truncate distally, ca. 12 mm long, 3–5 mm wide.